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These shares were sold in multiple transactions at prices ranging from $81.12 to $81.435 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (3) on this Form 4."}],"balance_after":36035,"source":{"system":"sec-edgar","filing":"0000874015-25-000288","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":2,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000288/edgardoc.xml","form_type":"4"}},{"id":982643,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-12-04T00:01:07.000Z","date":"2025-12-01","kind":"sale","planned":true,"acquired":false,"shares":20571,"price":80.6651,"value":1659361.7721,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 26, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F2","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $80.11 to $81.11 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (2) on this Form 4."}],"balance_after":43464,"source":{"system":"sec-edgar","filing":"0000874015-25-000288","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":1,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000288/edgardoc.xml","form_type":"4"}},{"id":993139,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Non-Qualified Stock Option (right to buy)","published_at":"2025-11-20T22:12:23.000Z","date":"2025-11-18","kind":"exercise","planned":false,"acquired":false,"shares":16000,"price":0,"value":0.0,"ownership":{"direct":true},"balance_after":0,"derivative":{"exercise_price":24.42,"exercise_date":"2017-07-01","expiration_date":"2026-06-30","underlying":"Common Stock","underlying_shares":16000},"source":{"system":"sec-edgar","filing":"0000874015-25-000282","document":"edgardoc.xml","document_sequence":1,"table":"derivative","row_type":"transaction","position":3,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000282/edgardoc.xml","form_type":"4"}},{"id":993138,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-11-20T22:12:23.000Z","date":"2025-11-18","kind":"exercise","planned":false,"acquired":true,"shares":16000,"price":24.42,"value":390720.0,"ownership":{"direct":true},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Acquired as a result of exercising a stock option that was scheduled to expire on 6/30/2026."}],"balance_after":16000,"source":{"system":"sec-edgar","filing":"0000874015-25-000282","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":1,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000282/edgardoc.xml","form_type":"4"}},{"id":1010162,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-11-04T01:14:56.000Z","date":"2025-10-31","kind":"sale","planned":true,"acquired":false,"shares":11200,"price":75.1586,"value":841776.32,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 26, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F5","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $75.00 to $75.41 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (5) on this Form 4."}],"balance_after":64035,"source":{"system":"sec-edgar","filing":"0000874015-25-000272","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":4,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000272/edgardoc.xml","form_type":"4"}},{"id":1010161,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-11-04T01:14:56.000Z","date":"2025-10-30","kind":"sale","planned":true,"acquired":false,"shares":5975,"price":73.85,"value":441253.75,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 26, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F4","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $73.6750 to $74.28  inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (4) on this Form 4."}],"balance_after":75235,"source":{"system":"sec-edgar","filing":"0000874015-25-000272","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":3,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000272/edgardoc.xml","form_type":"4"}},{"id":1010160,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-11-04T01:14:56.000Z","date":"2025-10-30","kind":"sale","planned":true,"acquired":false,"shares":8341,"price":73.16,"value":610227.56,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 26, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F3","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $72.63 to $73.63 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (3) on this Form 4."}],"balance_after":81210,"source":{"system":"sec-edgar","filing":"0000874015-25-000272","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":2,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000272/edgardoc.xml","form_type":"4"}},{"id":1010159,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-11-04T01:14:56.000Z","date":"2025-10-30","kind":"sale","planned":true,"acquired":false,"shares":2484,"price":72.19,"value":179319.96,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 26, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F2","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $71.62 to $72.62 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (2) on this Form 4."}],"balance_after":89551,"source":{"system":"sec-edgar","filing":"0000874015-25-000272","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":1,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000272/edgardoc.xml","form_type":"4"}},{"id":1028490,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-10-02T23:36:16.000Z","date":"2025-10-01","kind":"sale","planned":true,"acquired":false,"shares":16800,"price":65.4991,"value":1100384.88,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 26, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F2","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $65.02 to $65.86 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (2) on this Form 4."}],"balance_after":92035,"source":{"system":"sec-edgar","filing":"0000874015-25-000246","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":1,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000246/edgardoc.xml","form_type":"4"}},{"id":1048787,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-09-05T00:40:10.000Z","date":"2025-09-03","kind":"sale","planned":true,"acquired":false,"shares":11200,"price":60.1383,"value":673548.96,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 16, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F4","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $60.00 to $60.53 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (4) on this Form 4."}],"balance_after":108835,"source":{"system":"sec-edgar","filing":"0000874015-25-000223","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":3,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000223/edgardoc.xml","form_type":"4"}},{"id":1048786,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-09-05T00:40:10.000Z","date":"2025-09-02","kind":"sale","planned":true,"acquired":false,"shares":5600,"price":57.5311,"value":322174.16,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 16, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F3","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $57.500 to $57.575 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (3) on this Form 4."}],"balance_after":120035,"source":{"system":"sec-edgar","filing":"0000874015-25-000223","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":2,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000223/edgardoc.xml","form_type":"4"}},{"id":1048785,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-09-05T00:40:10.000Z","date":"2025-09-02","kind":"sale","planned":true,"acquired":false,"shares":11200,"price":55.0386,"value":616432.32,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 16, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F2","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $55.000 to $55.180 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (2) on this Form 4."}],"balance_after":125635,"source":{"system":"sec-edgar","filing":"0000874015-25-000223","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":1,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000223/edgardoc.xml","form_type":"4"}},{"id":1048738,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-09-05T00:34:45.000Z","date":"2025-09-03","kind":"sale","planned":true,"acquired":false,"shares":11200,"price":60.1383,"value":673548.96,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 16, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F4","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $60.00 to $60.53 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (4) on this Form 4."}],"balance_after":108835,"source":{"system":"sec-edgar","filing":"0000874015-25-000216","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":3,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000216/edgardoc.xml","form_type":"4"}},{"id":1048737,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-09-05T00:34:45.000Z","date":"2025-09-02","kind":"sale","planned":true,"acquired":false,"shares":5600,"price":57.5311,"value":322174.16,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 16, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F3","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $57.500 to $57.575 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (3) on this Form 4."}],"balance_after":120035,"source":{"system":"sec-edgar","filing":"0000874015-25-000216","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":2,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000216/edgardoc.xml","form_type":"4"}},{"id":1048736,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Common Stock","published_at":"2025-09-05T00:34:45.000Z","date":"2025-09-02","kind":"sale","planned":true,"acquired":false,"shares":11200,"price":55.0386,"value":616432.32,"ownership":{"direct":false,"nature":"by Trust"},"footnotes":[{"field":"shares","element":"transactionShares","label":"F1","text":"Shares sold pursuant to a Rule 10b5-1 Trading Plan adopted by the reporting person on December 16, 2024."},{"field":"price","element":"transactionPricePerShare","label":"F2","text":"The price reported in Column 4 is a weighted average price. These shares were sold in multiple transactions at prices ranging from $55.000 to $55.180 inclusive.  The reporting person undertakes to provide to Ionis Pharmaceuticals, Inc. any security holder of Ionis Pharmaceuticals, Inc. or the staff of the Securities and Exchange Commission, upon request, full information regarding the number of shares sold at each separate price within the ranges set forth in footnote (2) on this Form 4."}],"balance_after":125635,"source":{"system":"sec-edgar","filing":"0000874015-25-000216","document":"edgardoc.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":1,"url":"https://www.sec.gov/Archives/edgar/data/874015/000087401525000216/edgardoc.xml","form_type":"4"}},{"id":1085974,"issuer":{"id":1182,"kind":"organization","name":"IONIS PHARMACEUTICALS INC","symbol":"IONS"},"owners":[{"id":72326,"kind":"person","name":"WENDER JOSEPH H","position":1,"cik":"0001073384","reported_name":"WENDER JOSEPH H","relationship":{"director":true,"officer":false,"ten_percent_owner":false,"other":false}}],"security":"Restricted Stock Unit","published_at":"2025-07-17T20:54:31.000Z","date":"2025-07-15","kind":"exercise","planned":false,"acquired":false,"shares":4079,"price":0,"value":0.0,"ownership":{"direct":true},"footnotes":[{"field":"derivative.exercise_price","element":"conversionOrExercisePrice","label":"F2","text":"Each Restricted Stock Unit represents a contingent right to receive one share of Ionis common stock, or its equivalent cash value."},{"field":"derivative.exercise_date","element":"exerciseDate","label":"F3","text":"Restricted Stock Units vest 100% on either the first anniversary of the date of grant or the next regularly scheduled annual meeting of stockholders of the Company, whichever occurs earlier, and are delivered to the reporting person on the July 15th following the vesting date. Upon vesting, the Restricted Stock Units will be paid in whole shares of Ionis common stock or cash as may be determined by the Company."},{"field":"derivative.expiration_date","element":"expirationDate","label":"F3","text":"Restricted Stock Units vest 100% on either the first anniversary of the date of grant or the next regularly scheduled annual meeting of stockholders of the Company, whichever occurs earlier, and are delivered to the reporting person on the July 15th following the vesting date. 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