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Each of Mr. Guy Levy, Soleus PE GP I, LLC and Soleus GP disclaims beneficial ownership of these securities held by Soleus PE and this report shall not be deemed an admission that they are the beneficial owners of such securities for purposes of Section 16 or for any other purpose, except to the extent of their respective pecuniary interests therein."}],"balance_after":0,"derivative":{"underlying":"Common Stock","underlying_shares":104608},"source":{"system":"sec-edgar","filing":"0001493152-20-009217","document":"ownership.xml","document_sequence":1,"table":"derivative","row_type":"transaction","position":7,"url":"https://www.sec.gov/Archives/edgar/data/1812305/000149315220009217/ownership.xml","form_type":"4"}},{"id":3147480,"issuer":{"id":46837,"kind":"organization","name":"Lyra Therapeutics, Inc.","symbol":"LYRA"},"owners":[{"id":144649,"kind":"organization","name":"Soleus Private Equity GP I, LLC","position":1,"cik":"0001812273","reported_name":"Soleus Private Equity GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144650,"kind":"organization","name":"Soleus Private Equity Fund I, L.P.","position":2,"cik":"0001752427","reported_name":"Soleus Private Equity Fund I, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144651,"kind":"organization","name":"Soleus PE GP I, LLC","position":3,"cik":"0001812274","reported_name":"Soleus PE GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21347,"kind":"organization","name":"Soleus Capital Master Fund, L.P.","position":4,"cik":"0001812275","reported_name":"Soleus Capital Master Fund, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21348,"kind":"organization","name":"Soleus Capital, LLC","position":5,"cik":"0001786724","reported_name":"Soleus Capital, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21349,"kind":"organization","name":"Soleus Capital Group, LLC","position":6,"cik":"0001812276","reported_name":"Soleus Capital Group, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21343,"kind":"person","name":"Levy Guy","position":7,"cik":"0001812305","reported_name":"Levy Guy","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}}],"security":"Series B Convertible Preferred Stock","published_at":"2020-05-15T22:16:57.000Z","date":"2020-05-05","kind":"exercise","acquired":false,"shares":11666666,"price":0,"value":0.0,"ownership":{"direct":false,"nature":"See footnote"},"footnotes":[{"field":"derivative.exercise_price","element":"conversionOrExercisePrice","label":"F4","text":"Each share of preferred stock was automatically converted into 0.0289998 shares of common stock upon the closing of the Issuer's initial public offering."},{"field":"derivative.exercise_date","element":"exerciseDate","label":"F4","text":"Each share of preferred stock was automatically converted into 0.0289998 shares of common stock upon the closing of the Issuer's initial public offering."},{"field":"derivative.expiration_date","element":"expirationDate","label":"F4","text":"Each share of preferred stock was automatically converted into 0.0289998 shares of common stock upon the closing of the Issuer's initial public offering."},{"field":"ownership.nature","element":"natureOfOwnership","label":"F1","text":"The securities reported in this row are owned directly by Soleus Private Equity Fund I, L.P. (\"Soleus PE\"). Soleus Private Equity GP I, LLC (\"Soleus GP\") is the sole general partner of Soleus PE. Soleus GP holds voting and dispositive power over the shares held by Soleus PE. Soleus PE GP I, LLC is the sole manager of Soleus GP. Mr. Guy Levy is the sole managing member of Soleus PE GP I, LLC. Each of Mr. Guy Levy, Soleus PE GP I, LLC and Soleus GP disclaims beneficial ownership of these securities held by Soleus PE and this report shall not be deemed an admission that they are the beneficial owners of such securities for purposes of Section 16 or for any other purpose, except to the extent of their respective pecuniary interests therein."}],"balance_after":0,"derivative":{"underlying":"Common Stock","underlying_shares":338330},"source":{"system":"sec-edgar","filing":"0001493152-20-009217","document":"ownership.xml","document_sequence":1,"table":"derivative","row_type":"transaction","position":6,"url":"https://www.sec.gov/Archives/edgar/data/1812305/000149315220009217/ownership.xml","form_type":"4"}},{"id":3147479,"issuer":{"id":46837,"kind":"organization","name":"Lyra Therapeutics, Inc.","symbol":"LYRA"},"owners":[{"id":144649,"kind":"organization","name":"Soleus Private Equity GP I, LLC","position":1,"cik":"0001812273","reported_name":"Soleus Private Equity GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144650,"kind":"organization","name":"Soleus Private Equity Fund I, L.P.","position":2,"cik":"0001752427","reported_name":"Soleus Private Equity Fund I, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144651,"kind":"organization","name":"Soleus PE GP I, LLC","position":3,"cik":"0001812274","reported_name":"Soleus PE GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21347,"kind":"organization","name":"Soleus Capital Master Fund, L.P.","position":4,"cik":"0001812275","reported_name":"Soleus Capital Master Fund, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21348,"kind":"organization","name":"Soleus Capital, LLC","position":5,"cik":"0001786724","reported_name":"Soleus Capital, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21349,"kind":"organization","name":"Soleus Capital Group, LLC","position":6,"cik":"0001812276","reported_name":"Soleus Capital Group, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21343,"kind":"person","name":"Levy Guy","position":7,"cik":"0001812305","reported_name":"Levy Guy","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}}],"security":"Common Stock","published_at":"2020-05-15T22:16:57.000Z","date":"2020-05-05","kind":"purchase","acquired":true,"shares":155000,"price":16,"value":2480000.0,"ownership":{"direct":false,"nature":"See footnote"},"footnotes":[{"field":"ownership.nature","element":"natureOfOwnership","label":"F2","text":"The securities reported in this row are held by Soleus Capital Master Fund, L.P. (\"Master Fund\"). Soleus Capital, LLC is the sole general partner of Master Fund and thus holds voting and dispositive power over the shares held by Master Fund. Soleus Capital Group, LLC is the sole managing member of Soleus Capital, LLC. Mr. Guy Levy is the sole managing member of Soleus Capital Group, LLC. Each Soleus Capital Group, LLC, Soleus Capital, LLC and Mr. Guy Levy disclaims beneficial ownership of these securities held by Master Fund and this report shall not be deemed an admission that they are the beneficial owners of such securities for purposes of Section 16 or for any other purpose, except to the extent of their respective pecuniary interests therein."}],"balance_after":155000,"source":{"system":"sec-edgar","filing":"0001493152-20-009217","document":"ownership.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":5,"url":"https://www.sec.gov/Archives/edgar/data/1812305/000149315220009217/ownership.xml","form_type":"4"}},{"id":3147478,"issuer":{"id":46837,"kind":"organization","name":"Lyra Therapeutics, Inc.","symbol":"LYRA"},"owners":[{"id":144649,"kind":"organization","name":"Soleus Private Equity GP I, LLC","position":1,"cik":"0001812273","reported_name":"Soleus Private Equity GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144650,"kind":"organization","name":"Soleus Private Equity Fund I, L.P.","position":2,"cik":"0001752427","reported_name":"Soleus Private Equity Fund I, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144651,"kind":"organization","name":"Soleus PE GP I, LLC","position":3,"cik":"0001812274","reported_name":"Soleus PE GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21347,"kind":"organization","name":"Soleus Capital Master Fund, L.P.","position":4,"cik":"0001812275","reported_name":"Soleus Capital Master Fund, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21348,"kind":"organization","name":"Soleus Capital, LLC","position":5,"cik":"0001786724","reported_name":"Soleus Capital, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21349,"kind":"organization","name":"Soleus Capital Group, LLC","position":6,"cik":"0001812276","reported_name":"Soleus Capital Group, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21343,"kind":"person","name":"Levy Guy","position":7,"cik":"0001812305","reported_name":"Levy Guy","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}}],"security":"Common Stock","published_at":"2020-05-15T22:16:57.000Z","date":"2020-05-05","kind":"sale","acquired":false,"shares":16909,"price":16,"value":270544.0,"ownership":{"direct":false,"nature":"See footnote"},"footnotes":[{"field":"ownership.nature","element":"natureOfOwnership","label":"F1","text":"The securities reported in this row are owned directly by Soleus Private Equity Fund I, L.P. (\"Soleus PE\"). Soleus Private Equity GP I, LLC (\"Soleus GP\") is the sole general partner of Soleus PE. Soleus GP holds voting and dispositive power over the shares held by Soleus PE. Soleus PE GP I, LLC is the sole manager of Soleus GP. Mr. Guy Levy is the sole managing member of Soleus PE GP I, LLC. Each of Mr. Guy Levy, Soleus PE GP I, LLC and Soleus GP disclaims beneficial ownership of these securities held by Soleus PE and this report shall not be deemed an admission that they are the beneficial owners of such securities for purposes of Section 16 or for any other purpose, except to the extent of their respective pecuniary interests therein."},{"field":"ownership.nature","element":"natureOfOwnership","label":"F3","text":"On May 5, 2020, Soleus PE exercised a warrant to purchase an aggregate of 31,382 shares of the Issuer's common stock for $16.00 a share, which represents the Issuer's initial public offering price. In connection with the automatic net exercise of the warrant immediately prior to the closing of the Issuer's initial public offering, the Issuer withheld 16,909 of the warrant shares to pay the exercise price and issued the remaining 14,473 shares of common stock to Soleus PE. The Issuer also paid $13.77 to Soleus PE in lieu of a fractional share."}],"balance_after":457421,"source":{"system":"sec-edgar","filing":"0001493152-20-009217","document":"ownership.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":4,"url":"https://www.sec.gov/Archives/edgar/data/1812305/000149315220009217/ownership.xml","form_type":"4"}},{"id":3147477,"issuer":{"id":46837,"kind":"organization","name":"Lyra Therapeutics, Inc.","symbol":"LYRA"},"owners":[{"id":144649,"kind":"organization","name":"Soleus Private Equity GP I, LLC","position":1,"cik":"0001812273","reported_name":"Soleus Private Equity GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144650,"kind":"organization","name":"Soleus Private Equity Fund I, L.P.","position":2,"cik":"0001752427","reported_name":"Soleus Private Equity Fund I, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144651,"kind":"organization","name":"Soleus PE GP I, LLC","position":3,"cik":"0001812274","reported_name":"Soleus PE GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21347,"kind":"organization","name":"Soleus Capital Master Fund, L.P.","position":4,"cik":"0001812275","reported_name":"Soleus Capital Master Fund, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21348,"kind":"organization","name":"Soleus Capital, LLC","position":5,"cik":"0001786724","reported_name":"Soleus Capital, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21349,"kind":"organization","name":"Soleus Capital Group, LLC","position":6,"cik":"0001812276","reported_name":"Soleus Capital Group, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21343,"kind":"person","name":"Levy Guy","position":7,"cik":"0001812305","reported_name":"Levy Guy","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}}],"security":"Common Stock","published_at":"2020-05-15T22:16:57.000Z","date":"2020-05-05","kind":"other","acquired":true,"shares":31382,"price":8.63,"value":270826.66,"ownership":{"direct":false,"nature":"See footnote"},"footnotes":[{"field":"ownership.nature","element":"natureOfOwnership","label":"F1","text":"The securities reported in this row are owned directly by Soleus Private Equity Fund I, L.P. (\"Soleus PE\"). Soleus Private Equity GP I, LLC (\"Soleus GP\") is the sole general partner of Soleus PE. Soleus GP holds voting and dispositive power over the shares held by Soleus PE. Soleus PE GP I, LLC is the sole manager of Soleus GP. Mr. Guy Levy is the sole managing member of Soleus PE GP I, LLC. Each of Mr. Guy Levy, Soleus PE GP I, LLC and Soleus GP disclaims beneficial ownership of these securities held by Soleus PE and this report shall not be deemed an admission that they are the beneficial owners of such securities for purposes of Section 16 or for any other purpose, except to the extent of their respective pecuniary interests therein."},{"field":"ownership.nature","element":"natureOfOwnership","label":"F3","text":"On May 5, 2020, Soleus PE exercised a warrant to purchase an aggregate of 31,382 shares of the Issuer's common stock for $16.00 a share, which represents the Issuer's initial public offering price. In connection with the automatic net exercise of the warrant immediately prior to the closing of the Issuer's initial public offering, the Issuer withheld 16,909 of the warrant shares to pay the exercise price and issued the remaining 14,473 shares of common stock to Soleus PE. The Issuer also paid $13.77 to Soleus PE in lieu of a fractional share."}],"balance_after":474330,"source":{"system":"sec-edgar","filing":"0001493152-20-009217","document":"ownership.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":3,"url":"https://www.sec.gov/Archives/edgar/data/1812305/000149315220009217/ownership.xml","form_type":"4"}},{"id":3147476,"issuer":{"id":46837,"kind":"organization","name":"Lyra Therapeutics, Inc.","symbol":"LYRA"},"owners":[{"id":144649,"kind":"organization","name":"Soleus Private Equity GP I, LLC","position":1,"cik":"0001812273","reported_name":"Soleus Private Equity GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144650,"kind":"organization","name":"Soleus Private Equity Fund I, L.P.","position":2,"cik":"0001752427","reported_name":"Soleus Private Equity Fund I, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144651,"kind":"organization","name":"Soleus PE GP I, LLC","position":3,"cik":"0001812274","reported_name":"Soleus PE GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21347,"kind":"organization","name":"Soleus Capital Master Fund, L.P.","position":4,"cik":"0001812275","reported_name":"Soleus Capital Master Fund, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21348,"kind":"organization","name":"Soleus Capital, LLC","position":5,"cik":"0001786724","reported_name":"Soleus Capital, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21349,"kind":"organization","name":"Soleus Capital Group, LLC","position":6,"cik":"0001812276","reported_name":"Soleus Capital Group, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21343,"kind":"person","name":"Levy Guy","position":7,"cik":"0001812305","reported_name":"Levy Guy","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}}],"security":"Common Stock","published_at":"2020-05-15T22:16:57.000Z","date":"2020-05-05","kind":"exercise","acquired":true,"shares":104608,"price":0,"value":0.0,"ownership":{"direct":false,"nature":"See footnote"},"footnotes":[{"field":"ownership.nature","element":"natureOfOwnership","label":"F1","text":"The securities reported in this row are owned directly by Soleus Private Equity Fund I, L.P. (\"Soleus PE\"). Soleus Private Equity GP I, LLC (\"Soleus GP\") is the sole general partner of Soleus PE. Soleus GP holds voting and dispositive power over the shares held by Soleus PE. Soleus PE GP I, LLC is the sole manager of Soleus GP. Mr. Guy Levy is the sole managing member of Soleus PE GP I, LLC. Each of Mr. Guy Levy, Soleus PE GP I, LLC and Soleus GP disclaims beneficial ownership of these securities held by Soleus PE and this report shall not be deemed an admission that they are the beneficial owners of such securities for purposes of Section 16 or for any other purpose, except to the extent of their respective pecuniary interests therein."}],"balance_after":442948,"source":{"system":"sec-edgar","filing":"0001493152-20-009217","document":"ownership.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":2,"url":"https://www.sec.gov/Archives/edgar/data/1812305/000149315220009217/ownership.xml","form_type":"4"}},{"id":3147475,"issuer":{"id":46837,"kind":"organization","name":"Lyra Therapeutics, Inc.","symbol":"LYRA"},"owners":[{"id":144649,"kind":"organization","name":"Soleus Private Equity GP I, LLC","position":1,"cik":"0001812273","reported_name":"Soleus Private Equity GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144650,"kind":"organization","name":"Soleus Private Equity Fund I, L.P.","position":2,"cik":"0001752427","reported_name":"Soleus Private Equity Fund I, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":144651,"kind":"organization","name":"Soleus PE GP I, LLC","position":3,"cik":"0001812274","reported_name":"Soleus PE GP I, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21347,"kind":"organization","name":"Soleus Capital Master Fund, L.P.","position":4,"cik":"0001812275","reported_name":"Soleus Capital Master Fund, L.P.","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21348,"kind":"organization","name":"Soleus Capital, LLC","position":5,"cik":"0001786724","reported_name":"Soleus Capital, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21349,"kind":"organization","name":"Soleus Capital Group, LLC","position":6,"cik":"0001812276","reported_name":"Soleus Capital Group, LLC","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}},{"id":21343,"kind":"person","name":"Levy Guy","position":7,"cik":"0001812305","reported_name":"Levy Guy","relationship":{"director":false,"officer":false,"ten_percent_owner":true,"other":false}}],"security":"Common Stock","published_at":"2020-05-15T22:16:57.000Z","date":"2020-05-05","kind":"exercise","acquired":true,"shares":338330,"price":0,"value":0.0,"ownership":{"direct":false,"nature":"See footnote"},"footnotes":[{"field":"ownership.nature","element":"natureOfOwnership","label":"F1","text":"The securities reported in this row are owned directly by Soleus Private Equity Fund I, L.P. (\"Soleus PE\"). Soleus Private Equity GP I, LLC (\"Soleus GP\") is the sole general partner of Soleus PE. Soleus GP holds voting and dispositive power over the shares held by Soleus PE. Soleus PE GP I, LLC is the sole manager of Soleus GP. Mr. Guy Levy is the sole managing member of Soleus PE GP I, LLC. Each of Mr. Guy Levy, Soleus PE GP I, LLC and Soleus GP disclaims beneficial ownership of these securities held by Soleus PE and this report shall not be deemed an admission that they are the beneficial owners of such securities for purposes of Section 16 or for any other purpose, except to the extent of their respective pecuniary interests therein."}],"balance_after":338330,"source":{"system":"sec-edgar","filing":"0001493152-20-009217","document":"ownership.xml","document_sequence":1,"table":"nonDerivative","row_type":"transaction","position":1,"url":"https://www.sec.gov/Archives/edgar/data/1812305/000149315220009217/ownership.xml","form_type":"4"}}]}